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No set answer with a niche directory, as the total volume of applicable sites will vary depending upon how popular your niche is.
There wouldn't be a set answer anyway, it just depends on how you want to do it. It's usually never too early for the bots, the sooner you are in the index the better. Laying the trails for bots to visit means that human visitors could too, so you want it to look presentable, but that is a personal judgement call...
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